• Earl S. Tupper Fellow, Smithsonian Tropical Research Institute, Panama, 2015
• Sibbald Fellow, Royal Botanic Garden, Edinburgh, Scotland, 2015
• Formation postdoctorale. Institute of Systematic Botany-Munich Botanic Garden, l'Allemagne, 2014.
• Ph. D. en biologie évolutive, University of Connecticut, USA, 2011.
• M. Sc. en biologie des plantes, Southern Illinois University, USA, 2005.
• B. Sc. en biologie, Universidad de Panama, Panama, 2003

La liste complète se trouve sur le ResearchGate et Google Citations.

• Villarreal, J.C., Crandall-Stotler, B.J., Hart, M.L., Long, D.G. & Forrest, L.L. 2015. Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiopsida) with a slow molecular rate. New Phytologist. doi: 10.1111/nph.13716
• Villarreal, J.C. & K. Renzaglia. 2015. The hornworts: important advancements in early land plant evolution. Journal of Bryology 37(3): 157-170 (invited review).
• Brown, R.C., B.E. Lemmon, M. Shimamura, J.C. Villarreal & K.S. Renzaglia. 2015. Spores of relictual bryophytes: diverse adaptations to life on land. Review of Palaeobotany and Palinology 216: 1-17.
• Wicket, N., --42 authors, D.W. Stevenson, J.C. Villarreal, J.A. Leebens-Mack. 2014. A phylotranscriptomics analysis of the origin and early diversification of land plants. Proceedings of the National Academy of Sciences, USA.
• Li, F.-W., J.C. Villarreal, --32 authors, D.W. Stevenson,  K.M. Pryer. 2014. Horizontal gene transfer of a chimeric photoreceptor, neochrome, from bryophytes to ferns. Proceedings of the National Academy of Sciences, USA 111:  6672-6677,.
• Villarreal, J.C. & S.S. Renner. 2013. Correlates of monoicy and dioicy in hornworts, the apparent sister group to vascular plants. BMC Evolutionary Biology 13: 239. doi:10.1186/1471-2148-13-239.
• Villarreal, J.C. & K.S. Renzaglia. 2006. Structure and development of Nostoc strands in Leiosporoceros dussii (Anthocerotophyta): a novel symbiosis in land plants. American Journal of Botany 93: 693-705.

• BIO-1005 Diversité et écologie des végétaux

• Chercheur (Fellow), Smithsonian Tropical Research Institute, Panama
• Chercheur associé, Royal Botanic Garden Edinburgh, Scotland
• Membre de IBIS